Using gland regions on their heads, warthogs mark poles and other objects for self-orientation. Males often mark during the mating season. The male also sprays urine when searching for and inspecting sleeping cavities.
1958
1947
2008
2011
2017
1936
1978
2018
2022
1922
1999
1998
1992
2002
1993
1997
1996
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2001
1989